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The Chemistry of WineFrom Blossom to Beverage and Beyond$
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David R. Dalton

Print publication date: 2018

Print ISBN-13: 9780190687199

Published to Oxford Scholarship Online: November 2020

DOI: 10.1093/oso/9780190687199.001.0001

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PRINTED FROM OXFORD SCHOLARSHIP ONLINE (oxford.universitypressscholarship.com). (c) Copyright Oxford University Press, 2021. All Rights Reserved. An individual user may print out a PDF of a single chapter of a monograph in OSO for personal use. date: 26 October 2021

Roots, Shoots, Leaves, and Grapes

Roots, Shoots, Leaves, and Grapes

7 (p.22) (p.23) Roots, Shoots, Leaves, and Grapes
The Chemistry of Wine

David R. Dalton

Oxford University Press

As noted earlier and as anticipated by Charles and Francis Darwin it has been argued that plants sense the direction of gravity (gravitropism) by movement of starch granules found in cells called statocytes that contain compartments (organelles) called statoliths. The synthesis of statoliths appears to occur in the plastid (plant organelle) compartments called amyloplasts (Figure 7.1, 1). It has been suggested that this gravitropic signal then leads to movement of plant hormones such as indole-3-acetic acid (auxin) (Figure 7.2), through the phloem opposite to the pull of gravity to promote stem growth. Chloroplasts (Figure 7.1, 2) are cell compartments (plastids or organelles) in which photosynthesis is carried out. The process of photosynthesis, discussed more fully later, is accompanied by the production of adenosine triphosphate (ATP) from adenosine diphosphate (ADP) and inorganic phosphate (Pi) (Figure 7.3). ATP is consumed and converted to ADP and Pi in living systems. The cycle of production and consumption allows ATP to serve as an “energy currency” to pay for the reactions in living systems. Beyond this generally recognized critical function of chloroplasts, it has recently been pointed out that light/dark conditions affect alternative splicing of genes which may be necessary for proper plant responses to varying light conditions. The organelles or plastids which contain the pigments for photosynthesis and the amyloplasts that store starch are only two of many kinds of plastids. Other plastids, leucoplasts for example, hold the enzymes for the synthesis of terpenes, and elaioplasts store fatty acids. Apparently, all plastids are derived from proplastids which are present in the pluripotent apical and root meristem cells. The cell wall (Figure 7.1, 3) is the tough, rigid layer that surrounds cells. It is located on the outside of the flexible cell membrane, thus adding fixed structure. A representation of a portion of the cell wall (as made up of cellulose and peptide cross-linking) is shown below in Figure 7.7. The cells will have different sizes as a function of where they are found (e.g., leaf, stalk, root), but in every case, the cell wall limits the size of the membrane that lies within.

Keywords:   Arabidopsis thaliana, Darwin, Charles, Darwin, Francis, Golgi bodies, Vitis vinifera, xx, adenosine diphosphate (ADP), adenosine triphosphate (ATP), amyloplasts, apical meristem, aquaporin, branches, cell membrane, cell wall, cellulose, chloroplasts, cholesterol, choline, chromophore, cotyledons, cryptochromes, cytoplasm, dicotyledons, embryogenesis, embryonic leaves, endoplasmic reticulum, enzymatic processes, flavin adenine dinucleotide (FAD), folic acid, genome, glucose, glycans, glycerol, glycolipids, hexadecanoic acid, hydrophobic, indole-3-acetic acid, irradiation, lateral root formation, leaves, lignin, lipid bilayer, lipids, liposome, meristems, micelle, microtubule, mitochondria, nucleolus, oleic acid, organelles, outside cell membrane, palmitic acid, phloem transport system, phosphate anion (Pi), phosphatidylcholine, phosphatidylethanolamine, phosphatidylinositol, phosphatidylserine, photoreception, photosynthesis, plant cell, plant-pollinator interactions, plasma membrane, structure of, plastids, polysaccharide chains, primordial cells, proteins, pterion, reduced flavin adenine dinucleotide (FADH2), ribosomes, root walls, saccharides, self-pollination, shoot apical meristems, sphingomylein, statocytes, statoliths, transcription, true leaves, vacuole, vascular tissue, young leaf primordium

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